This is a contractile tissue. There are skeletal,
smooth and cardiac muscles.
Skeletal (striated,
voluntary) muscle
This acts mainly on the bony skeleton or as a
diaphragm, but it is also found around the pharynx and larynx, and forms some sphincters.
It is composed of unbranched fibres of sarcoplasm limited by a membrane, the sarcolemma,
and contains many nuclei. Each fibre has a motor endplate and contains many
contractile units, the myofibrils, which have
alternating dark (A) and light (I) bands. A dark line
(the Z disc) crosses the middle of the I band. The bands of adjacent myofibrils
coincide, giving the muscle fibre its striated appearance.Each fibre is
enveloped and attached to its neighbour by a fibrous endomysium, and bundles of
fibres are enclosed by a fibrous perimysium. A muscle composed of many bundles
is
surrounded by an epimysium.The motor nerve supply of
the muscle comes from the anterior horn cells of the spinal cord and the motor
nuclei of the brainstem. The sensory supply arises in the more specialized spindles
and tendon organs as well as the simpler touch
and pain endings. Impulses from the sensory endings
pass into the posterior horn of the spinal cord.Skeletal muscles are formed by
voluntary fibres. The muscles are usually attached at each end to bone, by the
periosteum,either directly or through tendons and aponeuroses,and cross one or more joints.
Occasionally two muscles meet
at a common stretchable union known as a raphe. Muscles have a very rich blood
supply.
Muscle fibres are arranged either parallel
to the direction of the action (sartorius), or obliquely to it.
aponeurosis is a flat, thin tendinous expansion
providing a wide attachment, as seen in the abdominal
wall muscles. When a movement occurs at a joint, the muscles concerned
in producing it are known as the prime movers
or agonists and those opposing it as antagonists.
Muscles contracting to steady the joint across which movement is occurring are
known as synergists. A further type of action is known as the action of
paradox, in which a muscle gradually relaxes against the pull of gravity, e.g.
bending forwards produced by relaxing the back muscles.
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